Hi,
I think you are the one confused. Your camera is able to communicate with the AI Lens through the AI Lever on both the lens and camera. Therefore, it is able to deduce the shutter speed based on the aperture setting on the lens.
When you "closed" down your aperture using the DOF preview lever, the AI lever on the camera is not released. Therefore, the camera will think there is lesser light entering the camera, and hence it will indicate under expose.
With a non-AI lens, there is no AI Lever on the lens. Hence, it is not able to communicate with the camera. Whatever aperture you set on the lens is not communicate with the camera. Therefore, you will need to use the DOF preview lever to do the metering.
Hence, when using your AI Lens, you need not have to "close" down your aperture to do the metering.
Hope this answer your question and explain everything.